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Colts fire six assistants, including D-coordinator Greg Manusky

INDIANAPOLIS -- The Colts have fired six assistant coaches, including defensive coordinator Greg Manusky, the team announced Wednesday.

Manusky arrived in Indianapolis in 2012 with coach Chuck Pagano, but the Colts' defense consistently struggled in all four seasons. It ranked in the 20s in overall defense three times in the four seasons, including 26th this past year.

The unit's highest finish was 11th in 2014 despite not having linebacker Robert Mathis the entire season.

The Colts also fired safeties coach Roy Anderson, secondary coach Mike Gillhamer, head strength and conditioning coach Roger Marandino, tight ends coach Alfredo Roberts and running backs coach Charlie Williams.

The Colts fired offensive coordinator Pep Hamilton after Week 8.

"We are grateful for the contributions these coaches have made to the Colts during their time here," Pagano said in a statement released by the team. "Obviously these days are never easy. We wish all of these men the best in their future endeavors."